Two processes, only one of them reverses
Connective tissue converts to gelatin above about 160F, fast above 180F, and is done by 195-205F. Gelatin coats the fibres, and that coating is what reads as succulent.
That is what makes a roast tender: gelatin coating the fibres. A second process runs at the same time and has nothing to do with collagen. Muscle proteins tighten from about 120F and squeeze out water. This never stops and nothing puts the water back. That water is gone whether or not the collagen ever converts. A roast that reaches 205F and falls apart at a fork has answered the tenderness question and said nothing about moisture. They were never the same measurement.
What to do: judge tenderness with a fork, not a clock, and accept that a fully tender roast has already given up water it will not get back. That's not a mistake, it's the trade. The fix from there is the sauce: a good gravy re-covers dry meat far more reliably than trying to cook around the physics.